Cytosolic carboxypeptidase 1 (AGTPBP1)

Metallocarboxypeptidase that mediates deglutamylation of target proteins. Catalyzes the deglutamylation of polyglutamate side chains generated by post-translational polyglutamylation in proteins such as tubulins.

Also removes gene-encoded polyglutamates from the carboxy-terminus of target proteins like MYLK. Acts as a long-chain deglutamylase and specifically shortens long polyglutamate chains, although it's unable to remove the branching point glutamate, a process catalyzed by AGBL5/CCP5.